Iran has sent short-range ballistic missiles to Russia and the Kremlin "will likely use them within weeks in Ukraine", the US secretary of state has said.
Speaking during a visit to the UK, Antony Blinken said the development was "a threat to all of Europe" and showed how Iran's "destabilising influence reaches far beyond the Middle East".
He told a news conference: "Dozens of Russian military personnel have been trained in Iran to use the Fath-360 close range ballistic missile system, which has a maximum range of 75 miles.
"Russia has now received shipments of these ballistic missiles and will likely use them within weeks in Ukraine against Ukrainians.
"Russia has an array of its own ballistic missiles, but the supply of Iranian missiles enables Russia to use more of its arsenal for targets that are further from the front line."
British Foreign Secretary David Lammy said the supply of ballistic missiles from Iran to Russia is "definitely a significant escalation" and said the UK is "coordinating".
However, when asked if he would approve Ukraine using the Storm Shadow long-range missiles to strike targets inside Russia, Mr Lammy said he would not comment on "operational issues".
The US will be announcing further sanctions on Iran later today, Mr Blinken announced, including further measures against Iran Air.
He said other countries are also expected to announce fresh sanctions.
However, the Ukrainian persident's chief of staff said imposing sanctions on Iran for supplying Moscow with ballistic missiles was not enough, and allies should allow Kyiv to carry out strikes deep into Russia using Western weapons.
"We also need permission to use Western weapons against military targets on the territory of the Russian Federation, deliveries of longer-range missiles, and strengthening our air defences," the official, Andriy Yermak, said on Telegram.
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Mr Lammy and Mr Blinken revealed they will be travelling together to Kyiv later this week, which Mr Lammy said is the "first joint visit of this kind for well over a decade".
Mr Blinken said Russia and Iran's relationship is a "two-way street", with the Kremlin sharing technology with Iran, including on nuclear issues and space information.
"So, as Iran's destabilising activity spread, so does Russia, sowing even greater insecurity in their regions and around the world," he said.
He added the US has been sharing intelligence behind the discovery with allies and partners "and working together to ensure that there will be significant economic consequences for Tehran's actions".
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